女主角突然生病了，所以她的替角必须立即上台表演。
The lead actress got sick suddenly, so her ___understudy had to perform immediately.
这种罕见疾病的病因目前研究不足。
The causes of this rare disease are still ___understudied.
他在排练期间一直在学习主角的台词，作为替补演员。
He has been ___understudying the lead role's lines during rehearsals.
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